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Simon and Daniel Wales :

They were brought into the Rapture project near its very start to design and build significant parts of the city. Note- a majority of the city wasn't "cathedrals", but a system of modular prefabricated building sections (Ryan and the other investors didn't have endless money after all).

They were likely victims of the economic distress caused by the major decline of City Construction (them being architects and builders) around 1952 -- economy disruption much like the problems facing Prentis Mill (AE bankruptcy) and others.

The Masons Quarter (previously a showcase of the Architects' art and a residential district) was repurposed to 'entertainment', as an organized Red Light District (as Sirens Alley) with at least one of the brothers (Daniel) becoming a local proprietor and provider of related services. Shady operations like Fontaine's Little Sister Orphanage were also located there and at least one ADAM bootleg operation. Simon Wales found a new calling when he adopted much of Sophia Lamb's philosophy, and ran a hidden cult church in the area. Sofia Lamb apparently resided in Sirens Alley not far from that 'church', before she was exiled to Persephone for Sedition.

When the Civil War/Anarchists Rebellion came around, Sirens Alley business probably fell off more than a little. It is not clear how much fighting took place in that vicinity. Later, with Sofia Lamb's attaining authority over the district, there was further deterioration and careless occupation, evidenced by what the Player sees years later. Daniel and Simon had influence and authority for running the district.

The Wales' operations and all the people in the area were killed (intentionally flooded by Sofia Lamb) apparently to make a 'Mommy Dearest' point to the Big Daddy known as Delta, (who was traveling through to attempt to rescue Sofia's daughter from Sofia's maniacal plans). Delta kills Simon to get ahold of a security key to move forward in his mission. Delta may have killed Daniel Wales or not, and Daniel possibly escaped the destruction Sofia carried out.

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Design of Rapture

Achievements

Reputation

Demise - Delta 'terminated' (maybe) each of the Brothers - wouldn't it have been nice to not have to kill them to get that passkey (or whatever was needed) to advance in the game?? For the MMORPG recycling these characters, we can always resolrt to the Vita Chamber loophole.

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Wales Bros Buildings Leaked, But So Did Many of Frank Lloyd Wright's (in the Real World) :

Except that Wright's leaks would NOT drown you (his buildings weren't built under the Ocean).

“Daniel… you and I drafted Rapture's blueprint together… "Wales and Wales, Architects." Do you remember  ?” ― Simon Wales

Daniel Wales (Audio Diary) : "Blasted Simon. So Rapture leaks… cryin' about it's only gonna add to the surplus o' moisture, isn't it ? Nothin's to say it's our design, but Simon goes to see Doctor Lamb anyway -- beset by guilt 'cause we can't find another contract... And he comes out the other side, sayin' he's always wanted to take the orders -- he's "Father Simon Wales" now ! If I ever see that Sofia Lamb in the street I'm gonna give her the back o' me fuckin' hand, and a knot on the head besides ! "

Daniel Wales (Audio Diary) : "(Sigh) Fuck it. I'm proper drunk and feelin' introspective. Now, in the motherland, Simon and I designed cathedrals together. But I was the black ol' sheep back there, never believin' a word o' the Jesus bollocks meself. Then Andrew Ryan says he wants Wales an' Wales for Rapture -- a Cathedral with no God, befittin' the ascent o'man. Now I'm runnin' girls... and Simon thinks he's saving souls. Father Wales -- Ha ! Can ya believe I shared a womb with such a sap ! "

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"Cathedrals" in the plural ? Not many Cathedrals were built at once (and were usually decades-long projects), and the Great Depression and WW2 also slowed down the available work. Acclaim isn't usually given to "Also Rans" who may have merely submitted designs for proposed "Cathedrals".

The Wales Bros were Architects, not Deep Water Engineers. I don't think Ryan would leave it upto 'architects' for critical engineering details like structural watertight integrity, do you ? For 'under the ocean' building - I doubt there would be many experienced 'architects' for that.

Basic theme/styles/shapes coordinated across Rapture - fine. So the losing their jobs (further contracts) is a rather weak/contrived story detail. Simpler Explanation for their financial distress/unemployment : After a while there just wasn't alot of NEW architecture construction (and design work), and possibly further work was when many affluent people already had enough of their preferred 'style' (already all around them). Once Rapture went into Chaos (after Ryan died) there wouldn't be much 'architectural' work anyway and the brothers would need to seek other employment.

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